Small trees or shrubs; branchlets glabrous. Leaves paripinnate with winged rachis; leaflets up to 11-jugate, sessile, very narrowly oblong to oblong-elliptic, entire, glandular-punctate and shiny above, otherwise glabrous. Inflorescence a terminal or axillary paniculate thyrse. Flowers functionally male or female. Sepals 5, free almost to base, ciliolate, persistent. Petals 5, ovate, ciliolate. Disc lanate. Stamens 5, inserted between lobes of disc, sterile in female flowers. Ovary 2-locular; locules 1-ovulate; ovule pendent from apex of locule; pistillode present in male flowers. Fruit drupaceous, 1- or 2-locular, glabrous. Seeds 1 or 2, without arillode.
Species 3, Old World tropics; sthn trop. Afr. 1: Filicium decipiens (Wight & Arn.) Thwaites, Zimbabwe, Malawi, Mozambique.
Monoecious evergreen trees. Leaves pinnate; leaflets in 1-3 pairs, opposite or alternate, entire, glabrous, lowermost leaflets often reduced and sometimes stipule-like. Inflorescences terminal and axillary, paniculate or simple. Flowers regular, greenish yellow. Sepals (4)5, free, unequal, strongly imbricate. Petals (0-4)5, ovate, clawed, fringed with small hairs. Disc pentagonal, lobed, convex. Stamens 5-7, hairy at base. Ovary 2(3)-locular and -lobed with 1 ovule per locule; style entire; ovary rudiment in male flowers a cushion of hairs. Fruit of 1(-3) indehiscent mericarps, developed ones ellipsoid, but 1 or 2 abortive at base. Seeds without aril.
Small trees or shrubs, usually single-stemmed and often with a palm-like habit, polygamous or dioecious. Leaves paripinnate, up to 1 m and longer, with +/- 4-8 pairs of +/- opposite leaflets; stipules absent. Inflorescences spicate racemes arranged in rows or groups on old wood or roots. Flowers slightly irregular with petals crowded to one side and stamens or ovary on opposite side. Calyx +/- urn-shaped, (4)5-lobed, finely, densely tomentose; lobes 1/4-1/2 as long as calyx. Petals 4 or 5(6), unequal, slightly longer than calyx, linear-spathulate, with a short scale and sometimes also a short subulate process above the claw. Disc annular, asymmetrical, lobed. Stamens 6-8(15); filaments filiform, conspicuously hairy; anthers 2-thecous, dorsifixed, linear-oblong. Ovary 3-9-locular and -lobed, with 1 erect ovule per locule; style present, sometimes arched in female flowers; stigma entire to slightly lobed. Fruit a large and fleshy, lobed or ribbed drupe. Seed 1 per locule, without aril.
Species 30 or more, trop. Africa; sthn trop. Afr. +/- 6, Angola (Cabinda).
Erect, glabrous shrubs, not or little branched. Leaves alternate, clustered near tips of branchlets, petiolate, persistent, elongate-obovate or -spathulate, apex rounded but tip abruptly long-acuminate, long-attenuate towards base, subcoriaceous, +/- sharply serrate-dentate; lateral nerves 14-18 pairs, prominent on both faces; stipules lanceolate, caducous. Inflorescence epiphyllous, 1-3(4)-flowered, on midrib of upper ?-3/4 of lamina, glabrous; median bract ovate-lanceolate, accrescent. Flowers bisexual or unisexual (plants polygamous), pedicelled. Sepals 3 or 4(5), imbricate, rather irregular and dry, ovate, ciliolate. Petals 4-6, imbricate, thin, much larger than sepals, ciliate. Stamens 25-40, free; filaments subulate; anthers almost basifixed, longitudinally dehiscent, apiculate by protruding connective. Ovary superior, glabrous, 1-locular, with 2-4 multi-ovulate parietal placentas; style simple, shortly 2-4-fid. Capsule opening with 2 or 3 valves; pericarp woody. Seeds weakly arillate, with endosperm.
Erect perennial herbs with simple or sparsely branched, glabrous, quadrangular stems. Leaves opposite, shortly petiolate, ovate to lanceolate, acuminate at apex, serrulate except cuneate base, scabrid above, penninerved. Inflorescence a lax raceme with a pair of flowers at each floriferous node. Flowers 2-lipped, reddish purple with paler throat. Calyx glabrous, 5-lobed; lobes ovate to lanceolate. Corolla slightly irregular; upper lip 2-lobed, with glandular hairs inside; lower lip 3-lobed; tube gradually widening, much longer than lobes. Stamens 4; filaments of anterior stamens geniculate and incurved, papillate and with glandular hairs, filaments of posterior stamens +/- straight, much shorter; anthers with 2 equal thecae. Ovary globose, glabrous; style filiform; stigma 2-lobed. Fruit a septicidal capsule. Seeds with alveolate endosperm.
Species 4, Old World tropics; sthn trop. Afr. 1: Artanema longifolium (L.) Vatke, Angola.
Trees; branches terete, striate. Leaves alternate, sessile, lanceolate, acuminate, base cuneate; stipules absent. Flowers bisexual or unisexual (dioecious), in fascicles on old wood or on branches, often subtended by spirally arranged bracts. Calyx a single whorl of 5 free, quincuncial sepals. Corolla broadly tubular; tube shorter than the 5 broadly ovate lobes. Stamens +/- 20 in a single whorl, +/- 3 opposite each corolla lobe and arising at top of corolla tube; filaments well-developed, glabrous; anthers extrorse, glabrous, absent in female flowers; staminodes 5 or more, much shorter than corolla lobes, usually petaloid, fimbriate, glabrous, often inflexed and covering ovary. Disc absent. Ovary many-locular; placentation axile; style usually included. Fruit usually depressed-globose; several- to many-seeded; outer pericarp often woody. Seeds strongly laterally compressed; testa smooth, shining; scar adaxial, nearly full-length, very narrow; embryo with thin foliaceous cotyledons and exserted radicle; endosperm copious.
Species +/- 6, W and WC trop. Africa; sthn trop. Afr. 1: Omphalocarpum mayumbense Greves, Angola (Cabinda).
Annual, rigid, erect, branched herbs. Leaves opposite to alternate, linear, entire or trifid. Flowers small, solitary-axillary or terminally racemose, bi-bracteolate. Calyx campanulate, 5-lobed; lobes ovate to lanceolate, acute or obtuse. Corolla limb subequally 5-lobed; lobes entire; tube enlarged above, sometimes incurved. Stamens 4, somewhat didynamous, included; anthers free, bithecal; thecae arched or parallel, distinct, apically attached. Ovary 2-4-locular; ovules many. Capsule globose to subglobose. Seeds many, obovoid.
Species 5, tropical west, east and southern Africa with 1 in India; sthn trop. Afr. 1: Micrargeria filiformis (Schumach. & Thonn.) Hutch. & Dalz., Angola, Zambia, Zimbabwe, Malawi, Mozambique, and sthn Afr.
Small slender trees, spuriously polygamous, often monoecious; bark smooth, grey; branchlets fulvous-tomentose. Leaves paripinnate, shortly petiolate or subsessile; leaflets 3- or 4-jugate, oblong-elliptic, lowest pair often resembling stipules. Inflorescence a +/- terminal or axillary, paniculate or racemoid thyrse. Flowers regular, whitish, sweetly scented. Sepals 5, almost free. Petals 5, each with a short broad pilose or pubescent scale. Disc patelliform or collar-like with a free scarious margin. Stamens 8; staminodes shorter in female flowers. Ovary 3 locular, densely hairy, locules 1-ovulate; pistillode subglobose in male flowers. Fruit capsular, 3-gonous, 3-locular, loculicidally dehiscent; valves lanuginous at base within. Seeds 1 per locule, ellipsoid, with an arillode at base.
Species 10, trop. Africa; sthn trop. Afr. 4, Angola, Zambia.
Tall, deciduous, often dioecious trees, often with strongly buttressed or deeply fluted bole. Leaves usually spirally arranged, rarely opposite, chartaceous or coriaceous, +/- clearly pellucid-punctate, densely pubescent to subglabrous; lateral nerves many-paired, looped distally and running parallel to leaf margin or extending to leaf edge to form the thickened margin, tertiary veins conspicuous; stipules 0. Flowers often unisexual, (4)5(6)-merous, sessile or shortly pedicellate, fasciculate in axils of current or fallen leaves. Calyx a single whorl of 4-6 sepals +/- free to base, or 6-11 in a closely imbricate spiral. Corolla: tube slightly shorter to longer than lobes; lobes 4-6(-9), simple, sometimes fringed or papillose, usually erect. Stamens 4-6(-9), arising in lower or upper half of tube, rarely free; anthers usually extrorse or laterally dehiscent, included; staminodes, if present, usually as many as corolla lobes, small, +/- subulate or rarely petaloid. Disc present or absent. Ovary 1-6(-16)-locular, subglobose, densely pilose; placentation axile; style cylindric, slightly expanded towards apex, included or exserted. Fruit a subglobose to narrowly ellipsoid 1-several-seeded berry. Seed solitary, +/- ellipsoid; testa shining brown with conspicuous scar; endosperm present or absent; cotyledons fleshy.
Species +/- 200, Old and New World tropics, Africa +/- 5; sthn trop. Afr. 3, Angola, Zambia, Zimbabwe, Malawi, Mozambique.
Small trees or shrubs. Leaves paripinnate, 4-7-jugate, very large, usually crowded at ends of branches; leaflets petiolulate, oblong-elliptic, abruptly acute-acuminate, entire, glabrous. Inflorescences racemes, axillary or arising on old wood. Flowers regular; pedicels articulated near base. Calyx turbinate, tomentellous outside. Sepals 5, deltoid. Petals 0. Disc regular, cup-shaped or flat, smooth. Stamens 8 or 9; filaments twice geniculate in bud, pilose; anthers oblong, introrse, basi-dorsifixed. Ovary 3(4)-locular and 3(4)-lobed, with 1 ovule per locule; pistillode present in male flowers. Fruit indehiscent, 2- or 3(4)-lobed, turbinate, shortly stipitate; calyx and style +/- persistent. Seeds flattened, without aril.
Species +/- 15, mostly in W trop. Africa; sthn trop. Afr. 1: Placodiscus resendeanus Exell & Mendonca, Angola (Cabinda).