Berkheya

Perennial herbs or subshrubs, sometimes cobwebby or woolly. Leaves alternate or radical, rarely opposite, sometimes decurrent, subentire, toothed, pinnatifid or pinnatisect, teeth spinescent. Capitula radiate or discoid, several- to many-flowered, terminal on single stems or on branchlets, rarely axillary, rarely corymbose or umbellate. Involucre campanulate; bracts in few to many rows, connate at base, tips and often margins spiny. Receptacle flat, honeycombed fimbrilliferous. Ray florets neuter, in single row, strap-shaped. Disc florets bisexual, fertile or sometimes inner ones male, tubular, dilated above, deeply 5-lobed. Anthers linear, sagittate at base; with lanceolate, apical appendage. Style terete, undivided or with linear, obtuse branches. Cypselas +/- obovoid, ribbed, pilose or glandular, pubescent to glabrous. Pappus of short or long scales or bristles. x = 7, 8.