Trees or shrubs; branchlets glabrous. Leaves imparipinnate, 2-5(-8)-jugate; not crowded at tips of branches; leaflets +/- opposite, usually with pitted glands in upper (and sometimes in lower) surface; petiole and rachis not winged. Flowers in terminal and axillary thyrses. Calyx 2-5-lobed, or often bud rupturing irregularly to give a variable number of lobes. Petals 5, imbricate, puberulous. Stamens 10, 5 antepetalous ones somewhat shorter; filaments with hairy appendage at base. Disc annular to subcylindric, sometimes partially enveloping ovary, with furrows on outside accommodating stamens. Gynoecium of 5 carpels free, except for cohering styles, each carpel containing 1 ovule; styles very short, connate; stigmas 5. Fruits of 1-3(4) drupaceous mericarps. Seeds 1 per mericarp.
Species 35-40, pantropical; sthn trop. Afr. 5, Angola, Zambia.
Erect or sometimes scandent shrubs or small trees; bark of trunk and larger branches corky, pale brown, often with conical corky bosses surmounted, at least at first, by a prickle; smaller branches +/- prickly or unarmed, pubescent to +/- glabrous. Leaves not crowded at ends of branches, usually imparipinnate; leaflets opposite, in 2-7 pairs, usually asymmetric, margins crenate-serrate to crenate, sometimes entire; petioles and rachis winged. Inflorescences paniculate, axillary or terminal. Sepals 4 or 5(6), united at base. Petals 4 or 5(6), much longer than sepals, valvate, pubescent to glabrous outside. Stamens 8-10; filaments with a hairy appendage at base. Disc annular. Ovary 4- or 5-locular, globose or 4- or 5-lobed, with 1 pendulous ovule per locule; styles completely fused; stigma capitate, 4- or 5-lobed. Fruit a 4- or 5(6)-lobed, depressed-globose berry with 4 or 5(6) seeds.
Species 3 or 4, in trop. Africa, SE Asia and tropical Australia; sthn trop. Afr. 1: Harrisonia abyssinica Oliv., Angola, Zambia, Malawi, Mozambique.
Shrubs or small trees; at least younger parts +/- ferrugineous-pubescent or tomentose. Leaves often +/- crowded towards ends of branches, usually imparipinnate; leaflets opposite, in 3-5 pairs, entire to shallowly repand, base asymmetric. Inflorescence axillary, spiciform or in spiciform panicles, polygamous. Flowers usually in distant small glomerules. Sepals 4 or 5, united towards base, imbricate. Petals 4 or 5, free, imbricate. Stamens 4 or 5, arising beneath and between lobes of disc, reduced in female flowers. Disc fleshy, (3)4(5)-lobed. Gynoecium of 4 or 5 carpels, free or joined at base, each with 1 pendulous ovule; absent or vestigial in male flowers; styles 4 or 5, free. Fruit of drupaceous mericarps developing from 1-4 carpels of each flower. Seed 1 per mericarp.
Species 8, Old World tropics; sthn trop. Afr. 1: Brucea antidysenterica J.F.Mill., Angola, Zambia, Malawi.