Pappea

Shrubs or trees, monoecious, producing male and then female flowers either on the same inflorescence or on separate inflorescences. Leaves simple, usually petiolate, oblong or obovate, entire or sometimes toothed, especially on coppice shoots. Flowers small, regular; in panicles or racemes. Sepals 5, small, connate to form a cupular or saucer-shaped calyx; lobes ovate. Petals 4-6, +/- as long as calyx, villous within. Disc small, annular. Stamens 8-10, arising within disc; filaments villous. Ovary 2- or 3-locular, 2- or 3-lobed, hirsute, with a solitary ovule in each locule; style short, thick; stigma simple. Fruit usually globose, velvety, green to brown, with 1 or 2 locules aborting; splitting to reveal a seed enclosed in a jelly-like, orange-red arillode. Seeds globose, black, shiny, with basal, circular depression.